There were several culprits in the Yankees’ 3-2 loss to Toronto on Sunday, from a lack of offense to a bad outing by Dellin Betances.

But there was also one more: Miguel Andujar’s defense.

The third baseman didn’t make any errors Sunday, but his shaky performance on defense throughout his rookie season — coupled with the expanded rosters in September — resulted in Aaron Boone pulling Andujar for a defensive replacement in the top of the eighth inning with the Yankees clinging to a one-run lead.

Adeiny Hechavarria was inserted at third base, and Andujar’s bat was taken out of the cleanup spot.

Betances ended up blowing the lead, and with the Yankees down a run going into the bottom of the ninth, Boone was forced to pinch-hit Neil Walker for Hechavarria against Ken Giles.

Walker, who has struggled lately, flied to right, and the Yankees didn’t score in the inning as they dropped their second straight to the Blue Jays.

“You know that going in,” Boone said of the possibility of Andujar’s spot in the order coming up again. “I think that’s the first time it’s come around to hurt us, if it did indeed hurt us.”

Andujar has been the Yankees’ most consistent hitter for much of the second half, which is why he’s moved to the middle of the lineup.

On Sunday, he went 0-for-3 with a strikeout and made a questionable play at third when he and Gary Sanchez collided on a foul pop by Yangervis Solarte in the fourth inning.

But his bat is never in doubt, as he proved again Saturday, when Andujar’s seventh-inning grand slam got the Yankees back into the game.

Boone said the strategy was based in large part on having added players on the roster, which allowed him to use Hechavarria at third then go to the switch-hitting Walker against righty Giles.

Clearly, though, the Yankees would have felt more comfortable with Andujar hitting to start the ninth, one swing from tying the game.

Andujar entered Sunday having gone 10-for-27 with three extra-base hits in his past 29 plate appearances, while Walker has just three hits in his past 23 at-bats, with no extra-base hits.

And the switch was hardly the only reason the Yankees fell again, since they also couldn’t score against Mark Leiter Jr. in the eighth after Leiter had given up 18 runs — 10 earned — in his previous 8 ¹/₃ innings over eight games.